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~2–1.6 billion years agoReputable sourceWell documented

The First Complex Cells

On the timeline · around ~2–1.6 billion years ago · ProterozoicArcheanProterozoicThe First Complex Cells2.5 Ga2.3 Ga2 Ga1.8 Ga1.5 Ga1.3 Ga1 Ga

What happened

Complex cells — eukaryotes, with a nucleus and internal compartments — evolved from simpler microbes. A key step was endosymbiosis: one cell engulfed a bacterium that became the mitochondrion, the powerhouse of the cell (and later, in plants, the chloroplast).

Why it matters

All plants, animals, fungi, and protists are built from eukaryotic cells; their origin made large, complex, multicellular life possible.

Sources