~2–1.6 billion years agoReputable sourceWell documented
The First Complex Cells
On the timeline · around ~2–1.6 billion years ago · Proterozoic
What happened
Complex cells — eukaryotes, with a nucleus and internal compartments — evolved from simpler microbes. A key step was endosymbiosis: one cell engulfed a bacterium that became the mitochondrion, the powerhouse of the cell (and later, in plants, the chloroplast).
Why it matters
All plants, animals, fungi, and protists are built from eukaryotic cells; their origin made large, complex, multicellular life possible.
Sources
- Understanding Evolution, UC Berkeley. How important is endosymbiosis? · Reputable source