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About 17 to 15 million years agoPeer-reviewedWell documented

Elephants trade a long jaw for a trunk

On the timeline · around About 17 to 15 million years ago · The Miocene GrasslandsThe Miocene GrasslandsElephants trade a long jaw for a trunk19 Ma18 Ma17 Ma16 Ma15 Ma14 Ma13 Ma

What happened

Early proboscideans, the elephant lineage, fed for millions of years using a long, shovel-like lower jaw rather than a dedicated trunk. A 2024 study of Miocene proboscidean skulls found that as grasslands spread and opened up new habitat, different lineages split into different feeding strategies: Platybelodon kept a shortened lower jaw paired with a strong, flexible trunk suited to cutting plants that grew upright, while Choerolophodon specialized instead in cropping low, horizontally spreading vegetation. Across the elephant lineage as a whole, the study found that the trunk gradually took over feeding entirely, and the long mandible early proboscideans had relied on disappeared.

Why it matters

The trunk's takeover let elephant ancestors follow the same spreading grasslands that were reshaping horses at the same time, but by inventing an entirely different tool: a single flexible, grasping organ instead of faster legs and tougher teeth. It is the innovation modern elephants, the largest living land animals, still depend on for almost everything they do.

How we know

Researchers compared skull and jaw shape across multiple Miocene proboscidean genera, correlating the shrinking mandible and changing muscle-attachment points directly with each species' inferred feeding style and habitat, linking the anatomical shift to the same grassland expansion documented independently in the horse fossil record.

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Related timelines

  • Human Evolution · The grassland world this timeline builds is the one early hominins stood up in next
Part of a timelineThe Age of Mammals5 events · The 60 million years between the dinosaurs and the first humans, when rat-sized survivors grew into whales, horses, and elephants across a world remade by spreading grasslands.View all →